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In fact, it leaves out some of the most obvious ones, like Remedy (the commercial choice for help desks for ages now) and TFS, used by zillions of Microsoft developers.

Indeed, in some ways, they aren't even "competing" products, because they are in a totally different class of their own. TFS' bug tracking is notable for its tight integration with source control and SharePoint, for example. BugZilla is regarded
as an absolutely awful piece of junk, and is about as barebones as it gets. Mantis is good.

J.Ja