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I went to both the Apple store and HP's website and configured a Mac Pro and an HP Z800 as closely as possible–processor, RAM, storage, and graphics–and made no other changes to the default configuration presented by the vendor. That produced the following results:

The Mac Pro, with dual 2.66 GB Xeon processors, 12 GB RAM, 1 TB 7200 RPM SATA HDD, and ATI Radion HD 5870 graphics was $5199.

The HP Z800, with dual 2.66 GB Xeon processors, 12 GB RAM, 1 TB 7200RPM SATA HDD, and NVidia Quadro 6000 graphics was $11,162.